Anatidae is the biological
family that includes the
ducks and most duck-like waterfowl, such as
geese and
swan. The
Magpie-goose is no longer considered to be part of the
Anatidae, but is placed in its own family Anseranatidae. These are
birds that are modified for swimming, floating on the
water surface, and in some cases diving in at least shallow
water.
They have webbed feet and bills which are flattened to a
greater or lesser extent. Their feathers are excellent at
shedding water due to special oils. Anatidae are remarkable
for being one of the few families of birds that possess a
penis; they are adapted for copulation on the water only
and care must be taken when breeding ducks or geese that a
pool is provided for this purpose as attempts to copulate on
dry land will often lead to injury of the drake's penis.
Duck, eider and goose feathers and down have long been
popular for bedspreads, pillows, sleeping bags and coats.
The members of this family also have long been used for
food.
While the status of the Anatidae as a family is
straightforward, and there is little debate about which
species properly belong to it, the relationships of the
different tribes and subfamilies within it are poorly
understood. The listing in the box at right should be
regarded simply one of several possible ways of organising
the many species within the Anatidae.
Previously divided into six subfamilies, recent
anatomical studies by Livezey (1986; A phylogenetic analysis
of recent Anseriform genera, Auk 103: 737-754) showed
that the Anatidae are better treated in nine subfamilies.
This classification has been followed by Madge & Burn:
Dendrocygninae: whistling ducks
One pantropical genus, of distinctive long-legged
goose-like birds:
Dendrocygna (whistling
ducks, 9 species)
Thalassorninae: White-backed Duck
One genus in Africa, most closely related to the
subfamily Dendrocygninae, though also showing convergent
similarities to the subfamily Oxyurinae:
Five to seven extant genera with 27 living species,
mainly cool temperate Northern Hemisphere but also some
Southern Hemisphere species, with the
swans in two genera (three genera in some
treatments), and the
geese in four genera (three genera in some
treatments):
Cereopsis (Cape Barren Goose, 1 species,
sometimes transferred to Tadorninae)
Cnemiornis (New Zealand Geese, extinct)
Stictonettinae: Freckled Duck
One genus in Australia, formerly included in the
Oxyurinae, but with anatomy suggesting a distinct
ancient lineage perhaps closest to the Anserinae:
Stictonetta (Freckled
Duck, 1 species)
Plectropterinae: Spur-winged Goose
One genus in Africa, formerly included in the
'perching ducks', but closer to the Tadorninae:
Plectropterus (Spur-winged
Goose, 1 species)
Coscoroba Swan. The hump beyond is a second
individual.
Tadorninae: shelducks, sheldgeese and steamer-ducks
This group of larger, often semi-terrestrial
waterfowl can be seen as intermediate between Anserinae
and Anatinae. Recent revision has resulted in the
inclusion of 10 extant genera with 23 living species
(one probably extinct) in this subfamily, mostly from
the Southern Hemisphere but a few in the Northern
Hemisphere:
The
dabbling duck group, of worldwide distribution, were
previously restricted to just one or two genera, but has
now been extended to include 8 extant genera and about
55 living species, including several genera formerly
known as the "perching ducks":
The moa-nalos, of which 4 species in 3 genera are
known to date, are a peculiar group of flightless,
extinct Anatidae from the Hawaiian Islands. Gigantic in
size and with massive bills, they were believed to be
geese, but have been shown to be in reality very closely
related to the genus Anas. They evolved to fill the
ecological niche of turtles, ungulates and other
megaherbivores.
The 16 species of diving ducks, of worldwide
distribution, in 3 genera; Marmaronetta was formerly
included with the dabbling ducks but is now treated
here, and phylogenetic analysis of the probably extinct
Pink-headed Duck of India, previously treated
separately in Rhodonessa, has shown that it is
possibly better placed in Netta:
Marmaronetta (Marbled Duck, 1 species)
Netta (Red-crested Pochard and allies, 4 species,
one probably extinct)
Aythya (pochards, scaups, etc, 12 species, one
probably extinct)
Red-crested Pochard
Merginae: eiders, scoters, sawbills and other sea-ducks
There are 10 extant genera and 20 living species
(one or two extinct); most of this group occur in the
Northern Hemisphere, but two Mergus in the
Southern Hemisphere: